Prof. Dr. Elvira Hörandl

Prof. Dr. Elvira Hörandl

Abteilung Systematik, Biodiversität und Evolution der Pflanzen mit Herbarium

Untere Karspüle 2
37073 Göttingen

Telefon:+49 551 39 7843
Fax:+49 551 39 22329
Sprechzeiten:on Wednesdays 12.00-1.00 pm
Foto Prof. Dr. Elvira Hörandl
Wintersemester 2018/19
About me

Scientific career

10/2011 - present Professor for Systematics, Biodiversity and Evolution of Plants  with Herbarium (W3), University of Göttingen; Head of the Department of Systematics, Biodiversity and Evolution of Plants.
12/2006 – 09/2011 Adjunct Associate Professor (Privatdozent) and research group leader at the University of Vienna
10/1999 – 12/2006 Associate editor position at the International Association of Plant Taxonomy. Research group leader at the University of Vienna.
01/1996 – 09/1999 Postdoctoral researcher (FWF, ÖAW and ÖNB Projects) and Lecturer at the University of Vienna.
01/1993 – 12/1995 Scientific Assistant at the Austrian Academy of Sciences (ÖAW); Coordination of the project “Flora of Austria”.
10/1987 – 12/1992 Research Assistant at the Institute for Botany, University of Vienna. Head: Prof. Manfred A. Fischer

Academic education and degrees

Habilitation: Botany, University of Vienna, 2006

Conferral of doctorate: Botany, University of Vienna, 1991, Thesis advisor: Prof. Manfred A. Fischer

Diploma in Biology and Earth Sciences, University of Vienna, 1988. Mag. rer. nat, Thesis advisor: Prof. Harald Niklfeld

Other scientific activities, awards and honours

Other scientific activities
2017 Symposium organizer at the BIOSYST EU conference in Gothenburg
2009 – 2013 Managing Committee Member of the EU COST Action Action FA0903: Harnessing plant reproduction for crop improvement
2006 – 2011 Member of the Committee for Electronic Publication for the International Code of Botanical Nomenclature (Melbourne Code)
2011 Symposium organizer at the XVIIIth International Botanical Congress in Melbourne
2006 Symposium organizer at the XVIIth International Botanical Congress in Vienna
1999 – 2012 Leader of FWF (Austrian Research Fund) and ÖAW (Austrian Academy of Science) projects
1993 – 2011 Lecturer at the University of Vienna
1994 – 2011 Lecturer at the University of Agriculture in Vienna

Editorial service
2011 – present Editorial Board PloS One (Plant Evolution)
2016 – 2018  Editor of Perspectives in Plant Ecology, Evolution and  Systematics
2006 – 2010 Editorial board member of TAXON
1999 – 2006 Associate editor of the journal TAXON (International Association for Plant Taxonomy)

Research interests and recent projects

I am interested in the evolution of asexual polyploid complexes in plants. The origin of apomixis (the asexual reproduction via seed) via hybridization and polyploidy is a main focus of my projects. Asexual genome evolution is shaped by rapid divergence of lineages and diversifying selection between sexual and apomictic lineages.

Furthermore, I want to explore the causality of the phenomenon that the geographical distribution of apomictic plants is usually much larger than their sexual relatives, especially in previously glaciated areas. Here we look at population genetic structure, epigenetic profiles modes of reproduction and ecology in plant species of the European Alps.

These studies have contributed to a new theory to resolve the "paradox of sex" in nature.Experimental approaches showed that apomixis and sexuality are responsive to environmental factors and support the hypothesis that sex is a response to physiological stress. Results suggest that DNA restoration during meiosis-mixis cycles provide the most important advantage to sexuality.

Hybridization among sexual species of willows (Salix) is being studied on alpine glacier forefields. The retreat of glaciers as a consequence of global warming and results in breakdown of ecological crossing barriers and upward migration of lowland plants. We study population genetic structure, ecology and fertility of willows to get first insights into evolutionary processes in the context of global warming. Furthermore we want to elucidate phylogenetic relationships of the genus Salix in Europe.

Phylogenetic analyses and and biogeographical studies of the large cosmopolitan genus Ranunculus (buttercups) are further topics of my research. Phylogenetic relationships, speciation processes and dispersal routes are being analyzed by using molecular phylogenies and molecular data. Currently we try to elucidate speciation processes in the Ranunculus auricomus complex. These studies are also the background for elaborating principles of an evolutionary classification.

Funded Projects on apomixis

  • The biodiversity of the apomictic Ranunculus auricomus polyploid complex, DFG project, 2017-2020, SPP 1991
  • The evolution of apomixis in autopolyploid alpine plants, DFG project, 2016-2019
  • The evolutionary establishment of apomixis in hybrids of the Ranunculus auricomus complex, DFG project, 2015-2018
  • The ecology of geographical parthenogenesis in alpine plants DFG-FWF project, 2013-2016
  • Reproductive strategies of weedy flowering plants in tropical rainforest transformation systems (DFG Project within SFB 990 - Tropical rainforest transofrmation systems), 2013-2015
  • Effects of hybridization on expression of apomixis in the Ranunculus auricomus complex (abstract), FWF-DFG project, 2009-2012
  • Geographical Parthenogenesis - a multidisciplinary study on alpine species (abstract), FWF project, 2006-2009
  • Apomixis in subnivale plants of the Alps, ÖAW project, 2009
  • Evolution of apomixis in the Ranunculus cassubicus complex (final report), FWF project, 2002-2006
  • Advantages of apomixis versus selfing (abstract), ÖAW project, 2006-2008
  • COST action Harnessing Plant Reproduction for Crop Improvement (HAPRECI), EU funded, 2009-2013 (pdf)
Funded projects on hybrid evolution and phylogeny of Salix
  • Reticulate evolution and hybridization in willows (Salix L.), DFG project, 2016-2019
  • Evolutionary processes on alpine glacier forefields: a pilot study on alpine willow hybrid populations. abstract (ÖAW 2010-2013)
  • A pilote study on hybrid speciation in alpine willows (Salix), (abstract), ÖAW 2008-2009)
  • Salix in Austria, 2002 (book)
Funded projects on the phylogeny of Ranunculus
  • Evolution, biogeographical history and ecology of high alpine and arctic species of Ranunculus (Ranunculaceae), 2011-2013
  • Character evolution, ecology, and a molecular phylogeny of Ranunculus s.l. (abstract), ÖAW and ÖAD projects, 2006-2009)
  • Phylogenetic analyses in the genus Ranunculus based on DNA sequences, ÖAW, 2002-2005,
  • Phylogeny and biogeographical history of buttercups (Ranunculus) in the Himalayas (abstract), National Geographic, 2008-2009)
Funded projects on the herbarium Göttingen
Wissenschaftliche und historische Erschließung des Herrenhäuser Herbariums in Göttingen, 2016-2018, Pro - Niedersachsen (MWK).


Selected publications

Journal papers


Hörandl, E. In press. The classification of asexual organisms: old myths, new facts, and a novel pluralistic approach. Taxon.

Barke, B.H., Daubert, M., Hörandl, E. 2018. Establishment of apomixis in diploid F2 hybrids and inheritance of apospory from F1 to F2 hybrids of the Ranunculus auricomus complex.  Frontiers in Plant Science 9:1111. doi: 10.3389/fpls.2018.01111               Open access.

Wagner, N.D., Gramlich, S., Hörandl, E. 2018. RAD sequencing resolved phylogenetic relationships in European shrub willows (Salix L. subg. Chamaetia and subg. Vetrix) and revealed multiple evolution of dwarf shrubs. Ecology & Evolution 8: 8243-8255. DOI: 10.1002/ece3.4360    Open access.

Hörandl, E., & Speijer, D. 2018. How Oxygen gave rise to eukaryotic sex. Proc. Roy. Soc. B 20172706. Open access.

Kirchheimer, B., Wessely, J., Gattringer, G., Hülber, K., Moser, D., Schinkel, C.C.F., Appelhans, M., Klatt, S., Caccianiga, M., Dellinger, A.S., Guisan, A., Kuttner, M., Lenoir, J., Maiorano, L., Nieto-Lugilde, D., Plutzar, C., Svenning, J.C., Willner, W., Hörandl, E., & Dullinger, S. 2018. Reconstructing geographical parthenogenesis: effects of niche differentiation and reproductive mode on Holocene range expansion of an alpine plant. Ecol. Letters 21: 392–401.   Open access.

Klatt, S., Schinkel, C.C.F., Kirchheimer, B., Dullinger, S., & Hörandl, E. 2018. Effects of cold treatments on fitness and mode of reproduction in the diploid and polyploid alpine plant Ranunculus kuepferi (Ranunculaceae). Annals of Botany 121: 1287–1298. DOI: 10.1093/aob/mcy017 Open access.


Schinkel CCF, Kirchheimer B, Dullinger S, Geelen D, De Storme N, Hörandl E. 2017. Pathways to polyploidy: indications of a female triploid bridge in the alpine species Ranunculus kuepferi (Ranunculaceae). Plant Syst. Evol.  303: 1093-1108. DOI 10.1007/s00606-017-1435-6. Open access.


Mirzaghaderi G, Hörandl E. 2016. The evolution of meiotic sex and its alternatives. Proc B  283: 20161221. doi: 10.1098/rspb.2016.1221 Open access.

Schinkel CCF, Kirchheimer B, Dellinger AS, Klatt S, Winkler M, Dullinger S, Hörandl E. 2016. Correlations of polyploidy and apomixis with elevation and associated environmental gradients in an alpine plant. AoB Plants. 10.1093/aobpla/plw064 Open access.

Kirchheimer B, Schinkel CF, Dellinger A, Klatt S, Moser D, Winkler M, Lenoir J , Caccianiga M, Guisan A, Nieto-Lugilde D, Svenning JC, Wilfried Thuiller W, Vittoz P, Willner W, Zimmermann NE, Hörandl E, Dullinger S. 2016. A matter of scale: Apparent niche differentiation of diploid and tetraploid plants may depend on extent and grain of analysis. Journal of Biogeography 43, 716–726   doi:10.1111/jbi.12663  Open access!

Gramlich S, Hörandl E. 2016. Fitness of natural willow hybrids in a pioneer mosaic hybrid zone. Ecology and Evolution 6: 7645-7655. DOI: 10.1002/ece3.2470. Open access.

Gramlich S, Sagmeister P, Dullinger S, Hadacek F, Hörandl E. 2016. Evolution in situ: Hybrid origin and establishment of willows (Salix L.) on alpine glacier forefields. Heredity 116, 531–541. doi:10.1038/hdy.2016.14   Open access!

Klatt S, Hadacek F, Hodac L, Brinkmann G, Eilerts M, Hojsgaard D, Hörandl E. 2016. Photoperiod extension enhances sexual megaspore formation and triggers metabolic reprogramming in  facultative apomictic Ranunculus auricomus. Frontiers in Plant Science 7: 278. doi: 10.3389/fpls.2016.00278.   Open access!

Baltisberger M, Hörandl E. 2016. Karyotype evolution supports the molecular phylogeny in the genus Ranunculus (Ranunculaceae). Persp. Plant Ecol., Evol. Syst. 18: 1-14.  doi:  Open access!

Dellinger A, Essl F, Hojsgaard D, Kirchheimer B, Klatt S, Dawson W, Pergl J, Pyšek, P, van Kleunen M, Weber E, Winter M, Hörandl E, Dullinger S. 2016. Niche dynamics of alien species do not differ among sexual and apomictic flowering plants. New Phytologist 209: 1313–1323.  Open access!

Clough Y, Krishna VV, Corre MD, Darras K, Denmead LH, Meijide A, Moser S, Musshoff O, Steinebach S, Veldkamp E, Allen K, Barnes AD, Breidenbach N, Brose U, Buchori D, Daniel R, Finkeldey R, Harahap I, Hertel D, Holtkamp AM, Hörandl E, Irawan B, Jaya INS, Jochum M, Klarner B, Knohl A, Kotowska MM, Krashevska V, Kreft H, Kurniawan S, Leuschner C, Maraun M, Melati DN, Opfermann N, Pérez-Cruzado C, Prabowo WE, Rembold K, Rizali A, Rubiana R, Schneider D, Tjitrosoedirdjo SS, Tjoa A, Tscharntke T, Scheu S 2016 Land-use choices follow profitability at the expense of ecological functions in Indonesian smallholder landscapes. Nature Communications 7, 13137 doi: 10.1038/ncomms13137.

Hodac L, Ulum FB, Opfermann N, Breidenbach N, Hojsgaard D,  Tjitrosoedirdjo SS, Vornam B, Finkeldey R, Hörandl E. 2016. Population genetic structure and reproductive strategy of the introduced Grass Centotheca lappacea in tropical land-use systems in Sumatra. PLoS ONE 11(1): e0147633. doi:10.1371/journal.pone.0147633. Open access!


Emadzade K, Lebmann M, Hoffmann MH, Tkach N, Lone F, Hörandl E (2015) Phylogenetic relationships and evolution of high mountain buttercups (Ranunculus) in North America and Central Asia. Perspect. Plant Ecol. Evol. Syst. 17: 131-141.

Hojsgaard D, Hörandl E (2015) A little bit of sex matters for genome evolution in asexual plants. Front. Plant Sci. 6:82. doi: 10.3389/fpls.2015.00082  Open access!


Hojsgaard D, Greilhuber J, Pellino M, Paun O, Sharbel TF, Hörandl E (2014) Emergence of apospory and bypass of meiosis via apomixis after sexual hybridisation and polyploidisation. New Phytologist 204: 1000-1012   DOI: 10.1111/nph.12954  Open Access!

Hodac L, Scheben AP, Hojsgaard D, Paun O, Hörandl E (2014) ITS Polymorphisms Shed Light on Hybrid Evolution in Apomictic Plants: A Case Study on the Ranunculus auricomus Complex. PLoS ONE 9(7): e103003. doi:10.1371/journal.pone.0103003. Open Access!

Hojsgaard D, Klatt S, Baier R, Carman JG, Hörandl E. 2014. Taxonomy and biogeography of apomixis in angiosperms and associated biodiversity characteristics. CRIT REV PLANT SCI. 33: 414-427. DOI: 10.1080/07352689.2014.898488   Open Access!

Hörandl E. (2014) Nothing in taxonomy makes sense except in the light of evolution: examples from the classification of Ranunculus. ANN MISS BOT GARDEN 100: 14-31.

Stuessy T., Hörandl E (2014). The importance of comprehensive phylogenetic (evolutionary) classification - a response to Schmidt-Lebuhn's commentary on paraphyletic taxa. CLADISTICS 30: 291-293.


Hörandl E. Hadacek F. (2013). The oxidative damage initiation hypothesis for meiosis. PLANT REPRODUCTION 26:351–367. doi: 10.1007/s00497-013-0234-7 Open access!

Pellino M, Hojsgaard D, Schmutzer T, Scholz U, Vogel H, Hörandl E, Sharbel TF (2013). Asexual genome evolution in the apomictic Ranunculus auricomus complex: examining the effects of hybridization and mutation accumulation. MOLEC ECOL 22: 5908–5921. doi: 10.1111/mec.12533

Cosendai AC, Wagner J, Ladinig U, Rosche C, Hörandl E (2013) Geographical parthenogenesis and population genetic structure in the alpine species Ranunculus kuepferi (Ranunculaceae). HEREDITY, 110: 560-569.


Hörandl E, Emadzade K (2012) Evolutionary classification: A case study on the diverse plant genus Ranunculus L. (Ranunculaceae). PERSPECT PLANT ECOL EVOL, 14: 310-324.

Hörandl E, Hojsgaard D (2012) The evolution of apomixis in Angiosperms: a reappraisal. PLANT BIOSYST, 146: 681-693.


Hörandl, E., Dobes, C., Suda, J., Vit, P., Urfus, T., Temsch, E. M., Cosendai, A.-C., Wagner, J., Ladinig, U. 2011. Apomixis is not prevalent in subnival to nival plants of the European Alps. Annals of Botany 108: 381–390.

Cosendai, A.-C., Rodewald, J., Hörandl, E. 2011. Origin and evolution of apomixis via autopolyploidy in the alpine plant species Ranunculus kuepferi. Taxon 60: 355-364.

Hörandl E., Emadzade K. 2011. The evolution and biogeography of alpine species in Ranunculus (Ranunculaceae) – a global comparison. Taxon 60: 415-426.

Hörandl, E. 2011. Evolution and biogeography of alpine apomictic plants. Taxon 60: 390-402.

Emadzade, K., Hörandl E. 2011. Northern Hemisphere origin, transoceanic dispersal, and diversification of Ranunculeae DC. (Ranunculaceae) in the Cenozoic. J. Biogeogr. 38: 517-530.

Emadzade, K., Gehrke, B., Linder, H.P., Hörandl, E. 2011. The biogeographical history of the cosmopolitan genus Ranunculus L. (Ranunculaceae) in the temperate to meridional zones. Molec. Phylog. Evol. 58: 4-21.


Hörandl, E., Stuessy, T.F. 2010. Paraphyletic groups as natural units of biological classification. Taxon 59: 1641-1653.

Emadzade, K., Lehnebach C., Lockhart P., Hörandl E. 2010. A molecular phylogeny, morphology and classification of genera of Ranunculeae (Ranunculaceae). Taxon 59: 809-828.

Hörandl E. 2010. Beyond cladistics: extending evolutionary classifications into deeper time levels. Taxon 59: 345-350.

Hörandl E. 2010. The evolution of self-fertility in apomictic plants. Sexual Pl. Repr. 23: 73-86.

Cosendai, A.-C., Hörandl, E. 2010. Cytotype stability, facultative apomixis and geographical parthenogenesis in Ranunculus kuepferi (Ranunculaceae). Ann. Bot. 105: 457-470.

Hoffmann MH., von Hagen, B, Hörandl E., Röser M., Tkach, N. 2010. Sources of the arctic flora: origins of arctic species in Ranunculus and related genera. Int. J. Pl. Sci. 171: 90–106.


Hörandl E. 2009. A combinational theory for maintenance of sex. Heredity 103: 445-457.

Hörandl, E., Temsch, E. 2009. Introgression of apomixis into sexual species is in the Ranunculus auricomus complex inhibited by mentor effects and ploidy barriers. Annals Bot 104: 81-89.

Hörandl, E, Greilhuber J., Klimova K., Paun O., Temsch E., Emadzade K & Hodálová I. 2009. Reticulate evolution and taxonomic concepts in the Ranunculus auricomus complex (Ranunculaceae): insights from morphological, karyological and molecular data. Taxon 58: 1194-1215.


Hörandl, E. 2008. Evolutionary implications of self-compatibility and reproductive fitness in the apomictic Ranunculus auricomus polyploid complex (Ranunculaceae). Int. J. Pl. Sci. 169: 1219-1228.

Hörandl, E., Cosendai, A.-C., Temsch, E. 2008. Understanding the geographic distributions of apomictic plants: a case for a pluralistic approach. Plant Ecology and Diversity 1: 309-320.


Hörandl, E. 2007: Neglecting evolution is bad taxonomy. Taxon 56: 1-5.

Lehnebach, C., Cano, A., Monsalve, C., McLenachan, P., Hörandl, E. & Lockhart, P. 2007: Phylogenetic relationships of the monotypic Peruvian genus Laccopetalum (Ranunculaceae) Pl. Syst. Evol. 264: 109-116.


Hörandl, E. 2006: Paraphyletic versus monophyletic taxa — evolutionary versus cladistic classifications. Taxon 55: 564–570.

Hörandl, E. 2006: The complex causality of geographical parthenogenesis. New Phytol. 171: 525–538.

Paun, O., Stuessy, T. F. & Hörandl, E. 2006: The role of hybridization, polyploidization and glaciation in the origin and evolution of the apomictic Ranunculus cassubicus complex. New Phytol. 171: 223–236.

Paun, O., Greilhuber, J., Temsch, E. & Hörandl, E. 2006: Patterns, sources and ecological implications of clonal diversity in apomictic Ranunculus carpaticola (Ranunculus auricomus complex, Ranunculaceae). Molec. Ecol. 15: 897–910.

Paun, O., Hörandl, E. 2006: Evolution of hypervariable microsatellites in apomictic polyploid lineages of Ranunculus carpaticola: directional bias at dinucleotide loci. Genetics 174: 387–398.


Hörandl, E., Paun, O., Johansson, J.T., Lehnebach, C., Armstrong, T., Chen, L., Lockhart, P. 2005: Phylogenetic relationships and evolutionary traits in Ranunculus s.l. (Ranunculaceae) inferred from ITS sequence analysis. Molec. Phyl. Evol. 36: 305-327.

Paun, O., Lehnebach, C., Johansson, J. T., Lockhart, P., Hörandl, E. 2005: Phylogenetic relationships and biogeography of Ranunculus and allied genera (Ranunculaceae) in the Mediterranean region and in the European Alpine System. Taxon 54: 911-930.

Books and book chapters

Hörandl E, Appelhans M (eds.) Next generation Sequencing in Plant Systematics. Regnum Vegetabile 158. IAPT, Bratislava.

Hojsgaard D, Hörandl E. 2015. Apomixis as a Facilitator of Range Expansion and Diversification in Plants. In: P.  Pontarotti (ed.), Evolutionary Biology: Biodiversification from Genotype to Phenotype. pp. 305-327. Springer.

Hörandl E. 2013. Meiosis and the paradox of sex in nature. In: Bernstein C, Bernstein, H. (ed) Meiosis. InTech, Rijeka, Croatia .

Hörandl E. 2009. Geographical parthenogenesis: opportunities for asexuality. In: Schön, I., Martens, K., Van Dijk P. (eds.), Lost Sex, pp. 161-186. Springer, Heidelberg.

Hörandl, press. Ranunculus. Systematische Gliederung. In: Gustav Hegi, Illustrierte Flora von Mitteleuropa, Bd. III Teil 3 (eds. Melzheimer V., Jäger E.). Jena, Weißdorn-Verlag.

Hörandl, E., Grossniklaus, U., van Dijk P. & Sharbel, T. (eds.), 2007. Apomixis: Evolution, Mechanisms and Perspectives. Gantner Verlag, Ruggell, Liechtenstein.

Hörandl, E. & Paun, O. 2007. Patterns and sources of genetic diversity in apomictic plants: implications for evolutionary potentials and ecology. In: Hörandl, E., Grossniklaus, U., Sharbel, T. & van Dijk, P. (eds.), Apomixis: Evolution, Mechanisms and Perspectives. Gantner Verlag, Ruggell, Liechtenstein.

Stuessy, T. F., Mayer, V., Hörandl, E. (eds.) 2003: Deep Morphology: Toward a Renaissance of Morphology in Plant Systematics. Gantner, Ruggel.

Hörandl, E., Florineth, F., Hadacek, F. 2002: Weiden in Österreich und angrenzenden Gebieten. Arbeitsbereich Ingenieurbiologie u. Landschaftsbau, Univ. Bodenkultur Wien, Wien. [Willows in Austria and adjacent regions. German with english abstract].

Stuessy, T. F., Hörandl, E., Mayer, V. (eds.) 2001: Plant Systematics: a Half-Century of Progress (1950-2000) and Future Challenges. IAPT, Vienna.

Invited talks
Salerno, Italy, Aug 2018. International Congress.
Paris, France, Feb 2018. International Congress.
San Diego, California, USA, Jan 2015. International Congress.
San Diego, California, USA, Jan 2013. International Congress.
Melbourne, Australia, Jul 2011. International Congress.
San Diego, California, USA, Jan 2011. International Congress.
Providence, Rhode Island, USA, July 2010. International Congress.
San Diego, California, USA. January 2010. International Congress.
Tübingen, Germany, June 2009. Symposium for Professor position.
Innsbruck, Austria, Dec 2008. Symposium for Professor position.
Tucson, Arizona, USA, Oct 2008. International Congress.
Zurich, Switzerland, Oct 2008. Seminar talk.
St Andrews, UK, June 2007. International Congress.
Berlin, Germany, June 2007. Symposium for Professor position.
Mainz, Germany, March 2007. International workshop.
Munich, Germany, April 2007. Seminar talk.
Zurich, Switzerland, Dec 2006. Seminar talk.
Vienna, Int. Bot. Congress, July 2005. International Congress.

Scientific Collaborations

University of Göttingen: Franz Hadacek, Stefan Scheu, Holger Kreft

International: Tim Sharbel (Saskatoon), Stefan Dullinger (Vienna), Maria Sartor (Argentina), Johann Greilhuber (Vienna), Johanna Wagner (Innsbruck), Danny Geelen (Ghent, Belgium), John Carman (Utah, USA), Tod F Stuessy (Ohio, USA), H. Peter Linder (Switzerland), Peter Lockhart (New Zealand), Jan Suda (Prague), Matthias Hoffmann (Halle, Germany), Matthias Baltisberger (Zurich), Ovidiu Paun (Vienna)

Scientific memberships: International Society for Plant Reproduction; International Association of Plant Taxonomy; Deutsche Botanische Gesellschaft; European Society of Evolutionary Biology; Botanical Society of America; German Society for Systematics (GfBS)

Scientific service: Editorial board member of the journal PLOS One. Editor of Perspectives in Plant Ecology, Evolution, and Systematics. Referee of the journals New Phytol, Evolution, Am J Bot, Molec Ecol, Int J Pl Sci, Annals Bot, Pl Syst Evol, J Biogeogr, Taxon, Syst Bot, and others. Reviewer panel of the US National Science Foundation, the German Science Foundation, and the Czech Science Foundation, the European Research Council, among others

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