Akademische Rätin / Akademischer Rat, Postdocs/ Senior lecturers   >   Dr. Dietrich Hertel, Akad. Oberrat
Dr. Dietrich Hertel, Akad. Oberrat

Phone: +49 551 39 5708
Fax: +49 551 39 22029
e-mail: dhertel@gwdg.de

Research interests

1) Ecophysiology of trees
2) Ecology and functioning of the tree root system
3) Forest ecosystems research (forest dynamics, biogeochemistry, climate change effects, functional biodiversity, land-use effects)
4) Ecology of the alpine treeline (global and regional aspects)

Current H Index (ISI web of science): 43
(updated 13-01-2023)

(for more details follow this link:)

Current H Index (Google Scholar Citations): 57
Current i10 Index (Google Scholar Citations): 119
(updated 13-01-2023)

(for more details follow this link:)

More details at ResearchGate:

Recent interdisciplinary research projects:




Curriculum vitae

1988-1994 Studies in Biology, Universities Karlsruhe and Göttingen

1995 Diploma in Biology, University of Göttingen

1995-1996 Scientific employee at the Centre for Forest Ecosystems Research, University of Göttingen

1999 Dr. rer. nat. (PhD) in Biology (Plant Ecology), University of Kassel

2000 Scientific employee at the Department of Plant Ecology and Ecosystems Research, University of Göttingen

2001-2017 Senior lecturer and researcher (Akademischer Rat) at the Department of Plant Ecology and Ecosystems Research, University of Göttingen

2002/2004/2005 Visiting scientist at CONICET-IANIGLA in Mendoza, Argentina (Instituto Argentino de Nivología, Glaciología y Ciencias Ambientales - National Institute for Snow, Glacier and Environmental Sciences Argentina)

Since 2017 Senior lecturer and researcher with higher rank (Akademischer Oberrat) at the Department of Plant Ecology and Ecosystems Research, University of Göttingen

Scientific activities and serviceship

Agricultural and Forest Meteorology
Agriculture, Ecosystems and Environment
Arid Land Research and Management
Applied Vegetation Science
Arctic, Antarctic and Alpine Research
Austral Ecology
Basic and Applied Ecology
Canadian Journal of Botany
Canadian Journal of Forest Research
Climate Research
Ecological Research
Ecology Letters
European Journal of Forest Research
Forest Ecology and Management
Frontiers in Plant Science

Global Change Biology
International Journal of Biometeorology
Journal of Biogeography
Journal of Ecology
Journal of Plant Nutrition and Soil Science
Journal of Tropical Ecology
Journal of Tropical Forest Science
Journal of Vegetation Science
Nature Communications
New Phytologist

Nutrient Cycling in Agroecosystems
Plant and Soil
Plant Ecology
Plant Ecology and Diversity
Tree Physiology



(for requesting a copy of an article, please contact me: dhertel(at)gwdg.de)

Peer-reviewed publications:

in press

Sierra Cornejo N, Becker JN, Hemp A, Hertel D (in press) Effects of land-use change and disturbance on the fine root biomass, dynamics, morphology, and related C and N fluxes to the soil of forest ecosystems at different elevations at Mt. Kilimanjaro (Tanzania). Oecologia.


Kotowska MM, Samhita S, Hertel D, Triadiati T, Beyer F, Allen K, Link R, Leuschner C (2023) Consequences of tropical rainforest conversion to tree plantations on fine root dynamics and functional traits. Oikos e08898.


Leuschner C, Feldmann E, Pichler V, Glatthorn J, Hertel D (2022) Forest management impact on soil organic carbon: A paired-plot study in primeval and managed European beech forests. Forest Ecology and Management 512: 120163.

Ramesha MN, Link RM, Paligi SS, Hertel D, Röll A, Hölscher D, Schuldt B (2022) Variability in growth-determining hydraulic wood and leaf traits in Melia dubia across a steep water availability gradient in southern India. Forest Ecology and Management 505: 119875.


Albrecht J, Peters MK, Becker JN, Behler C, Classen A, Ensslin A, Ferger SW, Gebert F, Gerschlauer F, Helbig-Bonitz M, Kindeketa WJ, Kühnel A, Mayr AV, Njovu HK, Pabst H, Pommer U, Röder J, Rutten G, Schellenberger Costa D, Sierra Cornejo N, Vogeler A, Vollstädt MGR, Dulle HI, Eardley CD, Howell KM, Keller A, Peters RS, Kakengi V, Hemp C, Zhang J, Manning P, Mueller T, Bogner C, Böhning-Gaese K, Brandl R, Hertel D, Huwe B, Kiese R, Kleyer M, Leuschner C, Kuzyakov Y, Nauss T, Tschapka M, Fischer M, Hemp A, Steffan-Dewenter I, Schleuning M (2021) Species richness is more important for ecosystem functioning than species turnover along an elevational gradient. Nature Ecology and Evolution 5: 1582–1593.

Röll A, Ramesha MN, Link RM, Hertel D, Schuldt B, Shekhargouda LP, Hölscher D (2021) Water availability controls the biomass increment of Melia dubia in South India. Forests 12: 1675.

Foerster A, Hertel D, Werner R, Leuschner C (2021) Belowground consequences of converting broadleaf to conifer forest: Comparing the fine root systems of European beech and Scots pine. Forest Ecology and Management 496: 119457.

Sierra Cornejo N, Leuschner, C, Becker JN, Hemp A, Schellenberger Costa D, Hertel D (2021) Climate implications on forest above- and belowground carbon allocation patterns along a tropical elevation gradient on Mt. Kilimanjaro (Tanzania). Oecologia 195: 797-812.

Kotowska MM, Link RM, Röll A, Hertel D, Hölscher D, Waite P-A, Moser G, Tjoa A, Leuschner C, Schuldt B (2021) Effects of wood hydraulic properties on water use and productivity of tropical rainforest trees. Frontiers in Forests and Global Change 3: 598759.


Sierra Cornejo N, Hertel D, Becker JN, Hemp A, Leuschner C (2020) Biomass, morphology and dynamics of the fine root system across a 3000 m-elevation gradient on Mt Kilimanjaro. Frontiers in Plant Science 11:13.

Mausolf K, Härdtle W, Hertel D, Leuschner C, Fichtner A (2020) Impacts of multiple environmental change drivers on growth of European beech (Fagus sylvatica): forest history matters. Ecosystems 23: 529-540.

Fuchs S, Hertel D, Schuldt B, Leuschner C (2020) Effects of summer drought on the fine root system of five broadleaf tree species along a precipitation gradient. Forests 11: 289.

Ahrens B, Guggenberger G, Rethemeyer J, John S, Marschner B, Heinze S, Angst G, Mueller CW, Kögel-Knabner I, Leuschner C, Hertel D, Bachmann J, Reichstein M, Schrumpf M (2020) Combination of energy limitation and sorption capacity explains 14C depth gradients. Soil Biology and Biochemistry 148: 107912.


Gundlach J, Krüger L, Herzberg C, Turdiev A, Poehlein A, Tascón A, Weiß M, Hertel D, Daniel R, Hänelt I, Lee VT, Stülke J (2019) Sustained sensing in potassium homeostasis: Cyclic di-AMP controls potassium uptake by KimA at the levels of expression and activity. Journal of Biological Chemistry 294: 2605-2614.

Kirfel K, Heinze S, Hertel D, Leuschner C (2019) Effects of bedrock type and soil chemistry on the fine roots of European beech - A study on the belowground plasticity of trees. Forest Ecology and Management 444: 256-268.

Peters MK, Hemp A, Appelhans T, Becker JN, Behler C, Classen A, Detsch F, Ensslin A, Ferger SW, Frederiksen SB, Gebert F, Gerschlauer F, Gütlein A, Helbig-Bonitz M, Hemp C, Kindeketa JW, Kühnel A, Mayr AV, Mwangomo E, Ngereza C, Njovu HK, Otte I, Pabst H, Renner M, Röder J, Rutten G, Schellenberger Costa D, Sierra-Cornejo N, Vollstädt MGR, Dulle HI, Eardley CD, Howe KM, Keller A, Peters RS, Ssymank R, Kakengi V, Zhang J, Bogner C, Böhning-Gaese K, Brandl R, Hertel D, Huwe B, Kiese R, Kleyer M, Kuzyakov Y, Nauss T, Schleuning M, Tschapka M, Fischer M, Steffan-Dewenter I (2019) Climate-land use interactions shape tropical mountain biodiversity and ecosystem functions. Nature 568: 88-92.

Röll A, Niua F, Meijide A, Ahongshangbam J, Ehbrecht M, Guillaume T, Gunawan D, Hardanto A, Hendrayanto, Hertel D, Kotowska MM, Kreft H, Kuzyakov Y, Leuschner C, Nomura M, Polle A, Rembold K, Sahner J, Hölscher D (2019) Transpiration on the rebound in lowland Sumatra. Agricultural and Forest Meteorology 274: 160-171.

Göbel L, Coners H, Hertel D, Willinghöfer S, Leuschner C (2019) The role of low soil temperature for photosynthesis and stomatal conductance of three graminoids from different elevations. Frontiers in Plant Science 10: 330.

Yang N, Butenschoen O, Grubert D, Rana R, Köhler L, Hertel D, Scheu S, Polle A, Pena R (2019) The effects of leaf litter type on biochemical composition of ectomycorrhizal fungi investigated by ATR-FTIR spectroscopy. Mycorrhiza 29: 85-96.


Guillaume T, Kotowska M, Hertel D, Knohl A, Krashevska V, Murtilaksono K, Scheu S, Kuzyakov Y (2018) Carbon costs and benefits of Indonesian rainforest conversion to plantations. Nature Communications 9: 2388.

Mausolf K, Härdtle W, Jansen K, Delory BM, Hertel D, Leuschner C, Temperton VM, von Oheimb G, Fichtner A (2018) Legacy effects of land-use modulate tree growth responses to climate extremes. Oecologia 187:825-837.

Mausolf K, Wilm P, Härdtle W, Jansen K, Schuldt B, Sturm K, von Oheimb G, Hertel D, Leuschner C, Fichtner A (2018) Higher drought sensitivity of radial growth of European beech in managed than in unmanaged forests. Science of the Total Environment 642: 1201-1208.

Angst G, Messinger J, Greiner M, Häusler W, Hertel D, Kirfel K, Kögel-Knabner I, Leuschner C, Rethemeyer J, Mueller CW (2018) Soil organic carbon stocks in topsoil and subsoil controlled by parent material, carbon input in the rhizosphere, and microbial-derived compounds. Soil Biology and Biochemistry 122: 19-30.

Redelstein R, Dinter T, Hertel D, Leuschner C (2018) Effects of inundation, nutrient availability and plant species diversity on fine root mass and morphology across a saltmarsh flooding gradient. Frontiers in Plant Science 9: 98.

Abou Rajab Y, Hölscher D, Leuschner C, Barus H, Tjoa A, Hertel D (2018) Effects of shade tree cover and diversity on root system structure and dynamics in cacao agroforests: the role of root competition and space partitioning. Plant and Soil 422: 349-369.

Meier, I, Knutzen F, Eder L, Müller-Haubold H, Goebel M-O, Bachmann J, Hertel D, Leuschner C (2018) The deep root system of Fagus sylvatica on sandy soil: structure and variation across a precipitation gradient. Ecosystems 21: 280-296.

Heinze S, Ludwig B, Piepho H-P, Mikutta R, Don A, Wordell-Dietrich P, Helfrich M, Hertel D, Leuschner C, Kirfel K, Kandeler E, Preusser S, Guggenberger G, Leinemann T, Marschner B (2018) Factors controlling the variability of organic matter in the top-and subsoil of a sandy Dystric Cambisol under beech forest. Geoderma 311: 37-44.


Allen K, Barnes AD, Kreft H, Corre MD, Jochum M, Veldkamp E, Clough Y, Daniel R, Darras K, Denmead LH, Haneda NF, Hertel D, Knohl A, Kotowska MM, Kurniawan S, Meijide A, Rembold K, Prabowo WE, Schneider D, Tscharntke T, Brose U (2017) Directing and cascading impacts of tropical land-use change on multitrophic biodiversity. Nature Ecology and Evolution 1: 1511-1519.

Becker J, Gütlein A, Sierra Cornejo N, Kiese R, Hertel D, Kuzyakov Y (2017) Legume and non-legume trees increase soil carbon sequestration in Savanna. Ecosystems 20: 989-999.

Kirfel K, Leuschner C, Hertel D, Schuldt B (2017) Root diameter and age are key determinants of the root hydraulic architecture of small- to medium-sized roots of mature beech trees in the top- and subsoil. Frontiers in Plant Science 8: 1194.

Gundlach J, Herzberg C, Hertel D, Thürmer A, Daniel R, Link H, Stülke J (2017) Adaptation of Bacillus subtilis to life at extreme potassium limitation. mBio 8: e00861-17.

Kubisch P, Leuschner C,  Coners H,  Gruber A, Hertel D (2017) Fine root abundance and dynamics of Stone pine (Pinus cembra) at the alpine treeline is not impaired by self-shading. Frontiers in Plant Science 8: 602.

Gundlach J, Herzberg C, Kaever V, Gunka K, Hoffmann T, Weiß M, Gibhardt J, Thürmer A, Hertel D, Daniel R, Bremer E, Commichau F M, Stülke J (2017) Control of potassium homeostasis is an essential function of the second messenger cyclic di-AMP in Bacillus subtilis. Science Signaling 10: eaal3011.

Homeier J, Báez S, Hertel D, Leuschner C (2017) Lessons learned from nutrient manipulation experiments in tropical forests. Frontiers in Earth Science 5: 27.

Gütlein A, Zistl-Schlingmann M, Becker JN, Sierra Cornejo N, Detsch F, Dannenmann M, Appelhans T, Hertel D, Kuzyakov Y, Kiese R (2017) Nitrogen turnover and greenhouse gas emissions in a tropical alpine ecosystem, Mt. Kilimanjaro, Tanzania. Plant and Soil 411: 243-259.


Clough Y, Krishna VV, Corre MC, Darras K, Denmead LH, Meijide A, Moser S, Musshoff O, Steinebach S, Veldkamp E, Allen K, Barnes A, Breidenbach N, Brose U, Buchori D, Daniel R, Finkeldey R, Harahap I, Hertel D, Holtkamp AM, Hörandl E, Irawan B, Jaya INS, Jochum M, Klarner B, Knohl A, Kotowska MM, Krashevska V, Kreft H, Kurniawan S, Leuschner C, Maraun M, Melati DN, Opfermann N, Pérez-Cruzado C, Prabowo WE, Rembold K, Rizali A, Rubiana R, Schneider D, Tjitrosoedirdjo SS, Tjoa A, Tscharntke T, Scheu S (2016) Land-use choices follow profitability at the expense of ecological functions in Indonesian smallholder landscapes. Nature Communications 7: 13137.

Meriem S, Tjitrosoedirjo S, Kotowska MM, Hertel D, Triadiati T (2016) Carbon and nitrogen stocks in dead wood of tropical lowland forests as dependent on wood decay stages and land-use intensity. Annals of Forest Research 59: 299-310.

Drescher J, Rembold K, Allen K, Beckschäfer P, Buchori D, Clough Y, Faust H, Fauzi AM, Gunawan D, Hertel D, Irawan B, Jaya INS, Klarner B, Kleinn C, Knohl A, Kotowska MM, Krashevska V, Krishna V, Leuschner C, Lorenz W, Meijide A, Melati D, Nomura M, Pérez-Cruzado C, Qaim , Siregar IZ, Steinebach S, Tjoa A, Tscharntke T, Wick B, Wiegand K, Kreft H, Scheu S (2016) Ecological and socioeconomic functions across tropical land-use systems after rainforest conversion. Philosophical Transactions of the Royal Society B 371:20150275.

Kubisch P, Hertel D. Leuschner C (2016) Fine root productivity and turnover of ectomycorrhizal and arbuscular mycorrhizal tree species in a temperate broad-leaved mixed forest. Frontiers in Plant Science 7: 1233.

Pransiska Y, Triadiati T, Soekisman T, Hertel D, Kotowska MK (2016) Forest conversion impacts on the fine and coarse root system, and soil organic matter in tropical lowlands of Sumatera (Indonesia). Forest Ecology and Management 379: 288-298.

Abou Rajab Y, Leuschner C, Barus H, Tjoa A, Hertel D (2016) Cacao cultivation under diverse shade tree cover allows high carbon storage and sequestration without yield losses. PLoS ONE 11: e0149949.

Kotowska MM, Leuschner C, Triadiati T, Hertel D (2016) Conversion of tropical lowland forest lowers nutrient return with litterfall, and alters nutrient use efficiency and seasonality of net primary production. Oecologia 180: 601-618.

Camenzind T, Homeier J, Dietrich K, Hempel S, Hertel D, Krohn A, Leuschner C, Oelmann Y, Olsson PA, Suárez JP, Rillig MC (2016) Opposing effects of nitrogen versus phosphorus additions on mycorrhizal fungal abundance along an elevational gradient in tropical montane forests. Soil Biology and Biochemistry: 94: 37-47.

Camenzind T, Papathanasiou HJ, Foerster A, Dietrich K, Hertel D, Homeier J, Oelmann Y, Olsson PA, Suárez JP, Rillig MC (2016) Increases in soil aggregation following phosphorus additions in a tropical premontane forest are not driven by root and arbuscular mycorrhizal fungal abundances. Frontiers in Earth Science 3:89.

Angst G, Kögel-Knabner I, Kirfel K, Hertel D, Mueller CW (2016) Spatial distribution and chemical composition of soil organic matter fractions in rhizosphere and non-rhizosphere soil under European beech (Fagus sylvatica L.). Geoderma 264: 179-187.


Kotowska MM, Leuschner C, Triadiati T, Meriem S, Hertel D (2015) Quantifying above-and belowground biomass carbon loss with forest conversion in tropical lowlands of Sumatra (Indonesia). Global Change Biology 21: 3620-3634.

Müller-Haubold H, Hertel D, Leuschner C (2015) Climatic drivers of mast fruiting in European beech and resulting C and N allocation shifts. Ecosystems 18: 1083-1100.

Kotowska MK, Hertel D, Abou Rajab Y, Barus H, Schuldt B (2015) Patterns in hydraulic architecture from roots to branches in six tropical tree species from cacao agroforestry and their relation to wood density and stem growth. Frontiers in Plant Science 6:191.

Chenlemuge T, Schuldt B, Dulamsuren C, Hertel D, Leuschner C, Hauck M (2015) Stem increment and hydraulic architecture of a boreal conifer (Larix sibirica) under contrasting macroclimates. Trees 29: 623-636.

Kubisch P, Hertel D. Leuschner C (2015) Do ectomycorrhizal and arbuscular mycorrhizal temperate tree species systematically differ in root order based fine root morphology and biomass? Frontiers in Plant Science 6: 64.

Chenlemuge T, Dulamsuren C, Hertel D, Schuldt B, Leuschner C, Hauck M (2015) Hydraulic properties and fine root mass of Larix sibirica along forest edge-interior gradients. Acta Oecologica 63: 28-35.

Violita, Kotowska MM, Hertel D, Triadiati, Miftahudin, Anas I (2015) Transformation of lowland rainforest into oil palm plantations results in changes of leaf litter production and decomposition in Sumatra, Indonesia. Journal of Biodiversity and Environmental Sciences 6: 546-556.

Scheibe A, Steffens C, Seven J, Jacob A, Hertel D, Leuschner C, Gleixner G (2015) Effects of tree species identity dominate over tree diversity on the soil microbial community structure. Soil Biology and Biochemistry 81: 219-227.

Köhler L, Leuschner, C, Hauck, M, Hertel D (2015) Cloud water interception and element deposition differ largely between Norway spruce stands along an elevation transect in Harz Mountains, Germany. Ecohydrology 8: 1048-1064.


Kessler M, Toivonen JM, Sylvester SP, Kluge J, Hertel D (2014) Elevational patterns of Polylepis tree height (Rosaceae) in the high Andes of Peru: role of human impact and climatic conditions. Frontiers in Plant Science: 5:194.

Hajek P, Leuschner C, Hertel D, Delzon S, Schuldt B (2014) Trade-offs between xylem hydraulic properties, wood anatomy and yield in Populus. Tree Physiology 34: 744-756.

Valtanen K, Eissfeller V, Beyer F, Hertel D, Scheu S, Polle A (2014) Carbon and nitrogen fluxes between beech and their ectomycorrhizal assemblage. Mycorrhiza 24: 645-650.

Jacob A, Hertel D, Leuschner C (2014) Diversity and species identity effects on fine root production and turnover in a species-rich temperate broad-leaved forest. Functional Plant Biology: 41: 648-689.

Moser G, Schuldt B, Hertel D, Horna V, Coners H, Barus H, Leuschner C (2014) Replicated throughfall exclusion experiment in an Indonesian perhumid rainforest: wood production, litter fall and fine root growth under simulated drought. Global Change Biology 20: 1481-1497.

Toivonen JM, Horna V, Kessler M, Ruokolainen K, Hertel D (2014) Interspecific variation in functional traits in relation to species climatic niche optima in Andean Polylepis (Rosaceae) tree species: evidence for climatic adaptations. Functional Plant Biology 41: 301-312.

Hajek P, Hertel D, Leuschner C (2014) Root order- and root age-dependent response of two poplar species to belowground competition. Plant and Soil 377: 337-355.

Leuschner C, Wulf M, Bäuchler P, Hertel D (2014) Forest continuity as a key determinant of soil carbon and nutrient storage in beech forests on sandy soils in northern Germany. Ecosystems 17: 497-511.


Hertel D, Strecker T, Müller-Haubold H, Leuschner C (2013) Fine root biomass and dynamics in beech forests across a precipitation gradient – is optimal resource partitioning theory applicable to water-limited mature trees? Journal of Ecology 101: 1183-1200.

Beyer F, Hertel D, Leuschner C (2013) Fine root morphological and functional traits in Fagus sylvatica and Fraxinus excelsior saplings as dependent on species, root order and competition. Plant and Soil 373: 143-156.

Müller-Haubold H, Hertel D, Seidel D, Knutzen F, Leuschner C (2013) Climate responses of aboveground productivity and allocation in Fagus sylvatica: a transect study in mature forests. Ecosystems 16: 1498-1516.

Hajek P, Hertel D, Leuschner C (2013) Intraspecific variation in root and leaf traits and leaf-root trait linkages in eight aspen demes (Populus tremula and P. tremuloides). Frontiers in Plant Science: 4:415.

Pfeiffer B, Fender A-C, Lasota S, Hertel D, Jungkunst HF, Daniel R (2013) Leaf litter is the main driver for changes in bacterial community structures in the rhizosphere of ash and beech. Applied Soil Ecology 72: 150-160.

Chenlemuge T, Hertel D, Dulamsuren C, Khishigjargal M, Leuschner C, Hauck M (2013) Extremely low fine root biomass in Larix sibirica forests at the southern drought limit of the boreal forest. Flora 208: 488-496.

Leuschner C, Wulf M, Bäuchler P, Hertel D (2013) Soil C and nutrient stores under Scots pine afforestations compared to ancient beech forests in the German Pleistocene: the role of tree species and forest history. Forest Ecology and Management 310: 405-415.

Leuschner C, Moser G, Hertel D, Erasmi S, Leitner D, Culmsee H, Schuldt B, Schwendenmann L (2013) Conversion of tropical moist forest into cacao agroforest: consequences for carbon pools and annual C sequestration. Agroforestry Systems 87: 1173-1187.

Eissfeller V, Beyer F, Valtanen K, Hertel D, Maraun M, Polle P, Scheu S (2013) Incorporation of plant carbon and microbial nitrogen into the rhizosphere food web of beech and ash. Soil Biology and Biochemistry 62: 76-81.

Rose L, Hertel D, Leuschner C (2013) Livestock type effects on biomass and nitrogen partitioning in temperate pastures with different functional group abundance. Grass and Forage Science 68: 386-394.

Leuschner C, Zach A, Moser G, Homeier J, Graefe S, Hertel D, Wittich B, Soethe N, Iost S, Röderstein M, Horna V, Wolf K (2013) The carbon balance of tropical mountain forests along an altitudinal transect. In: Bendix J, Beck E, Bräuning A, Makeschin F, Mosandl R, Scheu S, Wilcke W (Eds) Ecosystem Services, Biodiversity and Environmental Change in a Tropical Mountain Ecosystem of South Ecuador. Ecological Studies 221, Springer, Heidelberg, pp. 117-139.

Homeier J, Leuschner C, Bräuning A, Cumbicus NL, Hertel D, Martinson G, Spannl S, Veldkamp E (2013) Effects of nutrient addition on the productivity of montane forests and implications for the carbon cycle. In: Bendix J, Beck E, Bräuning A, Makeschin F, Mosandl R, Scheu S, Wilcke W (Eds) Ecosystem Services, Biodiversity and Environmental Change in a Tropical Mountain Ecosystem of South Ecuador. Ecological Studies 221, Springer, Heidelberg, pp. 315-329.

Beyer F, Hertel D, Jung K, Fender A-C, Leuschner C (2013) Competition effects on fine root survival of Fagus sylvatica and Fraxinus excelsior. Forest Ecology and Management 302: 14-22.

Cesarz S, Fender A-C, Beyer F, Valtanen K, Pfeiffer B, Gansert D, Hertel D, Polle A, Daniel R, Leuschner C, Scheu S (2013) Roots from beech (Fagus sylvatica L.) and ash (Fraxinus excelsior L.) differentially affect soil microorganisms and carbon dynamics. Soil Biology and Biochemistry 61: 23-32.

Jacob A, Hertel D, Leuschner C (2013) On the significance of belowground overyielding in temperate mixed forests: separating species identity and species diversity effects. Oikos 122: 463-473.

Rose L, Rubarth MC, Hertel D, Leuschner C (2013) Management alters interspecific leaf trait relationships and trait-based species rankings in permanent meadows. Journal of Vegetation Science 24: 239-250.


Kessler M, Hertel D, Jungkunst HF, Kluge J, Abrahamczyk S, Bos M, Buchori D, Gerold G, Gradstein SR, Köhler S, Leuschner C, Moser G, Pitopang R, Saleh S, Schulze CH, Sporn SG, Steffan-Dewenter I, Tjitrosoedirdjo SS, Tscharntke T (2012) Can joint carbon and biodiversity management in tropical agroforestry landscapes be optimized? PLoS ONE 7: e47192.

Homeier J, Hertel D, Camenzind T, Cumbicus NL, Maraun M, Martinson GO, Poma LN, Rillig MC, Sandmann D, Scheu S, Veldkamp E, Wilcke W, Wullaert H, Leuschner C (2012) Tropical Andean forests are highly susceptible to nutrient inputs - Rapid effects of experimental N and P addition to an Ecuadorian montane forest. PLoS ONE 7: e47128.


Seidel D, Beyer F, Hertel D, Fleck S, Leuschner C. (2011) 3D-laser scanning: a non-destructive method for studying above- ground biomass and growth of juvenile trees. Agricultural and Forest Meteorology 151: 1305-1311.

Toivonen JM, Kessler M, Ruokolainen K, Hertel D (2011) Accessibility predicts structural variation of Andean Polylepis forests. Biodiversity and Conservation 20: 1789-1802.

Hertel D, Köhler L, Rillig M (2011) Mycorrhizal, endophytic and ecomorphological status of tree roots in the canopy of a montane rain forest. Biotropica 43: 401-404.

Tscharntke T, Clough Y, Bhagwat SA, Buchori D, Faust H, Hertel D, Hölscher D, Juhrbandt J, Kessler M, Perfecto Y, Scherber C, Schroth G, Veldkamp E, Wanger TC (2011) Multifunctional shade-tree management in tropical agroforestry landscapes – a review. Journal of Applied Ecology 48: 619-629.

Graefe S, Leuschner C, Coners H, Hertel D (2011) Root functioning in tropical high-elevation forests: environmental vs. biological control of root water absorption. Environmental and Experimental Botany 71: 329-336.

Moser G, Leuschner C, Hertel D, Graefe S, Soethe N, Iost S (2011) Elevation effects on the carbon budget of tropical mountain forests (S Ecuador): the role of the belowground compartment. Global Change Biology 17: 2211-2226.

Hertel D, Schöling D (2011) Norway spruce shows contrasting changes in below- versus above-ground carbon partitioning towards the alpine treeline: evidence from a Central European case study. Arctic, Antarctic, and Alpine Research 43: 46-55.

Hertel D (2011) Tree roots in canopy soils of old European beech trees - an ecological reassessment of a forgotten phenomenon. Pedobiologia 54: 119-125.

Hertel D, Schöling D (2011) Below-ground response of Norway spruce to climate conditions at Mount Brocken (Germany) – a re-assessment of Central Europe's northernmost treeline. Flora 206: 127-135.


Hertel D, Leuschner C (2010) Fine root mass and fine root production in tropical moist forests as dependent on soil, climate and elevation. Chapter 47 in: Bruijnzeel LA, Scatena FN, Hamilton LS (Eds) Tropical montane cloud forests: Science for conservation and management. Cambridge University Press, Cambridge, UK, pp. 428-444.

Graefe S, Hertel D, Leuschner C (2010) N, P and K limitation of fine root growth along an elevation transect in tropical mountain forests. Acta Oecologica, 36: 537-542.

Schmidt-Lebuhn AN, Fuchs J, Hertel D, Hirsch H, Toivonen J, Kessler M (2010) An Andean radiation: Polyploidy in the tree genus Polylepis (Rosaceae, Sanguisorbeae). Plant Biology 12: 917-926.

Hertel D, Köhler L (2010) Are tree roots in the canopy ecologically important? A critical re-assessment from a case study in a tropical montane forest. Plant Ecology and Diversity 3: 141-150.

Moser G, Leuschner C, Hertel D, Hölscher D, Köhler M, Leitner D, Michalzik B, Prihastanti E, Tjitrosemito S, Schwendenmann L (2010) Response of cocoa trees (Theobroma cacao) to a 13-month desiccation period in Sulawesi, Indonesia. Agroforestry Systems 79: 171-187.

Schwendenmann L, Veldkamp E, Moser G, Hölscher D, Köhler M, Clough Y, Anas I, Djajakirana G, Erasmi S, Hertel D, Leitner D, Leuschner C, Michalzik B, Propastin P, Tjoa A, Tscharntke T, van Straaten O (2010) Effects of an experimental drought on the functioning of a cacao agroforestry system, Sulawesi, Indonesia. Global Change Biology 16: 1515-1530.

Moser G Leuschner C, Röderstein M, Graefe S, Soethe N, Hertel D (2010) Biomass and productivity of fine and coarse roots in five tropical mountain forests stands along an altitudinal transect in southern Ecuador. Plant Ecology and Diversity 3: 151-164.


Meinen C Hertel D, Leuschner C (2009) Root growth and recovery in temperate broad-leaved forest stands differing in tree species diversity. Ecosystems 12: 1103-1116.

Hertel D, Moser G, Culmsee H, Erasmi S, Horna V Schuldt B, Leuschner C (2009) Below- and above-ground biomass and net primary production in a paleotropical natural forest (Sulawesi, Indonesia) as compared to neotropical forests. Forest Ecology and Management 258: 1904-1912.

Meinen C, Leuschner C, Ryan NT, Hertel D (2009) No evidence of spatial root system segregation and elevated fine root biomass in multi-species temperate broad-leaved forests. Trees 23: 941-950.

Meinen C, Hertel D, Leuschner C (2009) Biomass and morphology of fine roots in temperate broad-leaved forests differing in tree species diversity: is there evidence of below-ground overyielding? Oecologia 161: 99-111.

Leuschner C, Harteveld M, Hertel D (2009) Consequences of increasing forest use intensity for biomass, morphology and growth of fine roots in a tropical moist forest on Sulawesi, Indonesia. Agriculture, Ecosystems, and Environment 129: 474-481.

Hertel D, Harteveld MA, Leuschner C (2009) Conversion of a tropical forest into agroforest alters the fine root-related carbon flux to the soil. Soil Biology and Biochemistry 41: 481-490.

Gaul D, Hertel D, Leuschner C (2009) Estimating fine root longevity in a temperate Norway spruce forest using three independent methods. Functional Plant Biology 36: 11-19.


Moser G, Röderstein M, Soethe N, Hertel D, Leuschner C (2008) Altitudinal changes in stand structure and biomass allocation of tropical mountain forests in relation to microclimate and soil chemistry. In: Beck E, Bendix J, Kottke I, Makeschin F, Mosandl R (Eds) Gradients in a Tropical Mountain Ecosystem of Ecuador. Ecological Studies 198, Springer, Berlin, Heidelberg, pp. 229-242.

Graefe S, Hertel D, Leuschner C (2008) Fine root dynamics along a 2000-m elevation transect in South Ecuadorian mountain rainforests. Plant and Soil 313: 155-166.

Hertel D, Therburg A, Villalba R (2008) Above- and below-ground response of Nothofagus pumilio to growth conditions at the transition from the steppe – forest boundary to the alpine treeline in southern Patagonia, Argentina. Plant Ecology and Diversity 1: 21-33.

Gaul D, Hertel D, Borken W, Matzner E, Leuschner C (2008) Effects of experimental drought on the fine root system of mature Norway spruce. Forest Ecology and Management 256: 1151-1159.

Graefe S, Hertel D, Leuschner C (2008) Estimating fine root turnover in tropical forests along an elevational transect using minirhizotrons. Biotropica 40: 536-542.

Gaul D, Hertel D, Leuschner C (2008) Effects of experimental frost on the fine root system of mature Norway spruce. Journal of Plant Nutrition and Soil Science 171: 690-698.

Srur AM, Villalba R, Villagra PE, Hertel D (2008) Influencias de las variaciones en el clima y en la concentración de C02 sobre el crecimiento de Nothofagus pumilio en la Patagonia. Revista chilena de historia natural 81: 239-256.

Hertel D, Wesche K (2008) Tropical moist Polylepis stands at the treeline in East Bolivia: the effect of elevation on stand microclimate, above- and below-ground structure, and regeneration. Trees 22: 303-315.


Moser G, Hertel D, Leuschner C (2007) Altitudinal change in LAI and stand leaf biomass in tropical montane forests: a transect study in Ecuador and a pan-tropical meta-analysis. Ecosystems 10: 924-935.

Harteveld M, Hertel D, Wiens M, Leuschner C (2007) Spatial and temporal variability of fine root abundance and growth in tropical moist forests and agroforestry systems (Sulawesi, Indonesia). Ecotropica 13: 111-120.

Hertel D, Leuschner C, Harteveld M, Wiens M (2007) Fine root mass, distribution and regeneration in disturbed primary forests and secondary forests of the moist tropics. In: Tscharntke T, Leuschner C, Zeller M, Guhardja E, Bidin A (Eds) Stability of tropical rainforest margins, linking ecological, economic and social constraints of land use and conservation. Springer, Berlin, pp. 89-108.

Leuschner C, Moser G, Bertsch C, Röderstein M, Hertel D (2007) Large elevational increase in tree root/shoot ratio in tropical mountain forests of Ecuador. Basic and Applied Ecology 8: 219-230.

Steffan-Dewenter I, Kessler M, Barkmann J, Bos MM, Buchori D, Erasmi S, Faust H, Gerold G, Glenk K, Gradstein SR, Guhardja E, Harteveld M, Hertel D, Höhn P, Kappas M, Köhler S, Leuschner C, Maertens M, Marggraf R, Migge-Kleian S, Mogea J, Pitopang R, Schaefer M, Schwarze S, Sporn SG, Steingrebe A, Tjitrosoedirdjo SS, Tjitrisoemito S, Twele A, Weber R, Woltmann L, Zeller M, Tscharntke T (2007) Tradeoffs between income, biodiversity, and ecosystem functioning during tropical rainforest conversion and agroforestry intensification. Proceeding of the National Academy of Science, USA (PNAS) 104: 4973-4978.


Hertel D, Leuschner C (2006) The in situ root chamber: a novel tool for the experimental analysis of root competition in forest soils. Pedobiologia 50: 217-224.

Leuschner C, Meier IC, Hertel D (2006) On the niche breadth of Fagus sylvatica: soil nutrient status in 50 Central European beech stands on a broad range of bedrock types. Annals of Forest Science 63: 355-368.

Hertel D, Hölscher D, Köhler L, Leuschner C (2006) Changes in fine root system size and structure during secondary succession in a Costa Rican montane oak forest. In: Kappelle M (ed) Ecology and Conservation of Neotropical Montane Oak Forests. Ecological Studies 185, Springer, Heidelberg. pp. 283-297.

Leuschner C, Wiens M, Harteveld M, Hertel D, Tjitrosemito S (2006) Patterns of fine root mass and distribution along a disturbance gradient in a tropical montane forest, Central Sulawesi (Indonesia). Plant and Soil 283: 163-174.


Röderstein M, Hertel D, Leuschner C (2005) Above- and below-ground litter production in three tropical montane forests in southern Ecuador. Journal of Tropical Ecology 21: 483-492.

Meier IC, Leuschner C, Hertel D (2005) Nutrient return with leaf litter fall in Fagus sylvatica forests across a soil fertility gradient. Plant Ecology 177: 99-112.


Leuschner C, Hertel D, Muhs A, Schmid I, Koch O, Hölscher D (2004) Stand fine root biomass and fine root morphology in old-growth beech forests as a function of precipitation and soil fertility. Plant and Soil 258: 43-56.


Leuschner C, Hertel D (2003) Fine root biomass of temperate forests in relation to soil acidity and fertility, climate, age and species. Progress in Botany 64: 405-438.

Hertel D, Leuschner C, Hölscher D (2003): Size and structure of fine root systems in old-growth and secondary tropical montane forests (Costa Rica). Biotropica 35: 143-153.

Bolte A, Hertel D, Ammer C, Schmid I, Nörr R, Kuhr M, Redde N (2003) Freilandmethoden zur Untersuchung von Baumwurzeln. Forstarchiv 74: 240-262.


Hertel D, Leuschner C (2002) A comparison of four different fine root production estimates with ecosystem carbon balance data in a Fagus-Quercus mixed forest. Plant and Soil 239: 237-251.

Hertel D, Leuschner C (2002) Erratum - A comparison of four different fine root production estimates with ecosystem carbon balance data in a Fagus-Quercus mixed forest. Plant and Soil 245: 335.

Hölscher D, Hertel D, Leuschner C, Hottkowitz M (2002) Tree species diversity and soil patchiness in a temperate broad-leaved forest with limited rooting space. Flora 197: 118-125.

Hölscher D, Hertel D, Koenies H (2002) Soil nutrient supply and biomass production in a mixed forest on a skeleton-rich soil and an adjacent beech forest. Journal of Plant Nutrition and Soil Science 165: 668-674.


Leuschner C, Hertel D, Coners H, Büttner V (2001) Root competition between beech and oak: a hypothesis. Oecologia 126: 276-284.

Leuschner C, Backes K, Hertel D, Schipka F, Schmitt U, Terborg O, Runge M (2001) Drought responses at leaf, stem and fine root levels of competitive Fagus sylvatica L. and Quercus petraea (Matt.) Liebl. trees in dry and wet years. Forest Ecology and Management 149: 33-46.


Hertel D, Leuschner C (1998) Die Rhizosphäre in einem Eichen-Buchenmischwald: Feinwurzelproduktion und die Bedeutung von Wurzelkonkurrenz. Verhandlungen der Gesellschaft für Ökologie 28: 441-447.

Leuschner C, Hertel D, Muhs A, Schmid I (1998) Feinwurzel-Bestandesmassen der Rotbuche an verschiedenen Standorten innerhalb ihrer ökologischen Amplitude in Nordwest- und Mitteldeutschland. Verhandlungen der Gesellschaft für Ökologie 28: 429-434.

Other publications:

Hertel D, Abou Rajab Y, Leuschner C (2016) Hoher Kakaoantrag auf Kosten der Umwelt – ein lösbarer Konflikt. Journal Culinaire 23: 34-41.

Hertel D, Schuldt B (2014) Holzanatomische Untersuchungen der hydraulischen Leitfähigkeit fossiler Birken und Kiefern der Jüngeren Dryas von Dissenchen 18. In: Schopper F (Hrg.): Ausgrabungen im Niederlausitzer Braunkohlerevier 2011/2012. Arbeitsberichte zur Bodendenkmalpflege in Brandenburg 27: 173-176.

Hertel D (2012) Einzigartig im Norden Mitteleuropas - Waldgrenze am Brocken ist natürlichen Ursprungs! Der Harz 03/2012: 13.

Hertel D (2012) Neue Forschungen zur Waldgrenze am Brocken. Neue Wernigeröder Zeitung 23: 9.

Hertel D (2012) Neue Forschungsergebnisse zur Waldgrenze am Brocken. Unser Harz 5: 98-99.

Meyer P, Schmidt M, Blick T, Brunet J, Dorow WHO, Hakes W, Härdtle W, Heinken T, Hertel D, Knapp HD, Leuschner C, von Oheimb G, Otte V, Schmidt W (2011) Stellungnahme zu Walentowski H. et al. 2010. Sind die deutschen Waldnaturschutzkonzepte adäquat für die Erhaltung der buchenwaldtypischen Flora und Fauna? Eine kritische Bewertung basierend auf der Herkunft der Waldarten des mitteleuropäischen Tief- und Hügellandes. (Forstarchiv 81: 195-217). Forstarchiv 82: 62-66.

Hertel D, Leuschner C (2008) Konkurrenz zwischen Waldbäumen – Interaktionsmechanismen im Kronen- und Wurzelraum unter besonderer Berücksichtigung der Buche. In: Nationalparkverwaltung Harz (Hrsg) Tun und Lassen im Naturschutz. Tagungsbericht zur 7. wissenschaftlichen Tagung im Nationalpark Harz. Schriftenreihe aus dem Nationalpark Harz 2: 23-28.

Hertel D, Coners H, Muhs A, Schipka F, Strobel J (2004) Zur Trockenheitsgrenze der Buche in Mittel- und Ostdeutschland: Eine ökosystemare Transektstudie an Buchen-Altbeständen. Berichte des Landesamtes für Umweltschutz Sachsen-Anhalt, Sonderheft 2/2004: 28-37.

Hertel D (1999) Das Feinwurzelsystem von Rein- und Mischbeständen der Rotbuche: Struktur, Dynamik und interspezifische Konkurrenz. Dissertationes Botanicae 317. Gebrüder Borntraeger Verlagsbuchhandlung Berlin, Stuttgart (Dissertation, Universität Kassel). 190 pp.

Leuschner C, Backes K, Hertel D, Schmitt U, Schipka F, Terborg O (1997) Wasserstress-Antwort auf Blatt-, Wurzel- und Stammebene von Rotbuchen und Traubeneichen in einem Altholz-Mischbestand in NW-Deutschland. Ecosys, Supplement 20: 11-24.

Hertel D (1995) Streuabbau in verschiedenen Stadien der Heide-Waldsukzession. Diplomarbeit (unveröffentlicht), Universität Göttingen. 106 pp.

Hertel D (1988) Orchis spitzelii Koch im Wallis - ein Neufund für die Schweiz. Mitteilungsblatt des Arbeitskreises Heimische Orchideen Baden-Württemberg 20: 883-889.

Hertel D (1988) Über das Vorkommen von Orchis spitzelii Koch im Wallis. Bullethin de la Murithienne 106: 75-78.




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